Demographic Transition

 

1.   The same forces that led people to go on crusades, and colonize the world, led to the growth of cities. Non-inheriting children went to cities to work as laborers and servants.

 

2.   Death rates exceeded birth rates in cities until the 19th century. However, the cities continued to grow because of migration. This and increased trade created a stimulus for a mercantile economy to grow and for technological innovation.

 

3.   Technological inovations occurred both in production (the industrial revolution) and in medical care and public health

 

4.   At the end of the 19th century the germ theory of disease started to impact mortality rates, and so did public health measures. In the last 5 years of the 19th century, many of the cities in Europe and America cleaned up the water supply, and this drastically reduced mortality rates, especially in the summer months.

 

5.   At the same time, fertility rates began to decrease as well.

 

6.   The decrease in mortality and fertility rates has been called the demographic transition. It has been described as a process in which 1) mortality rates decline 1st and populations grow; 2) people realize the decrease in mortality and respond by having smaller families. So fertility and mortality are back in balance and populations are stable. There have been many attempts to explain the transition, which happened in the developed world from about 1890-1920, and in the developing world after 1960 or so.

 

Demographic transition in Sweden and Mexico (mortality drops below fertility and populations grow, then they are equal, and afterwards, below replacement fertility occurs when fertility rates are lower than mortality rates; process is faster Mexico, because of modern medicine and modern economic conditions, but population grew faster as well.)

 

Possible Explanations

What is a theory of fertility?

A good theory should explain:

1)      cross-cultural variation in fertility

2)      within population variation in fertility

3)      secular trends in fertility

4)      relationships between relevant ecological variables and fertility outcomes (e.g. wealth, food intake, status)

 

Why the demographic transition? Doesn’t this contradict expectations based on evolutionary theory?

 

There are three main evolutionary hypotheses:

1)      quality-quantity trade-offs, economic costs of children, competitive environment – time frame of payoff?

2)      evolved psychology linked to sex or status, not reproduction per se, so current environment leads to maladaptive outcomes.

3)      cultural transmission – imitate successful individuals of low fertility, when individual learning costs are high.

 

Other theories

1)   Cultural equilibrium – Cultures respond to mortality so that values regarding reproduction serve to maintain a balance between births and deaths. There is a lag between the timing of mortality decline and the realization that fertility must also decline to avoid population growth.

 

Problems are:

 

1)   Group selection – why should individuals behave for the good of the group.

2)   The theory was used in the developing world with very poor results. The idea was that by improving medical care, people would have smaller families, but instead populations grew very fast.

3)   Fails to explain population growth with baby boom and below replacement fertility.

 

Trends in TFR and crude birth rates in US

Note baby boom and explain difference between crude birth rate (births/1000 in population), completed fertility (total births in a woman’s lifetime) and total fertility rate (synthetic expected completed fertility using the cross section).

 

Fertility decline in World regions

(note that fertility decline had already occurred in Europe and went to below replacement).

 

 

2) Caldwell’s wealth flows theory

 

Predemographic transition- Net flow of wealth is upwrd from offspring to parents. Children worked for the good of the kin group. Investment in kids when they are young, then kids give more back as they age. They also provide old-age insurance. When kids are a net asset, parents want as many as possible.

 

Post-demographic transition – kids are a net cost and parents only want a few of them – more like a consumption good. Kids are into their own selfish interests and do not give back. Emphasis on individualism as opposed to collectivism.

 

Contrast with evolutionary theory where net flows are always downward.

 

Problems:

1)   Logical – such a system is not possible because it would lead to runaway population growth.

2)   Emprical – Kaplan showed that even in traditional, predemographic transitions societies, like the Ache, children are a net cost; yet fertility is high and people really want lots of kids.

 

Evolutionary approach

Tradeoff between quantity and quality of offspring

Offspring Survival and Parental investment

Hypothetical interior optimum

Two-stage embodied capital model including offspring income

Multigenerational recursion

 

Male income as a function of sibling size from ABQ men

(there is a tradeoff between quantity and quality in terms of income, if not survival)

 

Fathers children on respondants children from ABQ men

(even if there is an effect of offspring income, there is no effect on fertility)

 

Anglo completed fertility

(no effect of income on fertility, but big effect of baby boom. Fertility is low before and after)

 

Hispanic completed fertility

(note the negative effect of income on fertiity in earlier cohorts. Perhaps there is a negative effect at early stage of demo transition when only people involved in the education-oriented part of the society, with higher incomes, have small families)

 

Number of children and grandchildren

(because income and fertility are unrelated, those that have the highest number of children have the most grandchildren, even if they have less income- no tradeoff)

 

Fertility and grandchildren

(bars show frequency distribution in percents of fertility, most people have 2 or 3 children, line shows linear relationship between fertility and fitness as measured by grandchildren- this is a big problem for evolutionary theory: Why is the most fit strategy not the most prevalent? Why did people respond to increased weath with less rather than more kids)

 

Two big problems (to be on exam)

1)   why is fertility so low

2)   why don’t wealthier people have more kids

 

 

 

People are living in a novel environment. Can we understand the modern response by understanding the proximate mechanisms that have evolved in the past to respond to environmental variation? We need a theory with two characteristics: 1) Predicts optimal responses under traditional conditions most similar to our evolutionary past; 2) predicts nonfitness maximizing response under modern circumstances.

 

 

Natural Selection and Fertility: The Problem

 

 

 

End part 1